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@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}*,*:before,*:after{transition:none!important}}
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@media(max-width:480px){.mentoring-section .contributor-list{grid-template-columns:1fr}.hero-figure .figure-index{display:none}}
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.project-index a,.jump-links a{padding-block:4px}
.contributor-list h3{font-size:15px;font-family:var(--sans);font-weight:500;letter-spacing:0;line-height:1.5}
.reset-search{margin-top:16px;border:1px solid #aebca5;color:var(--ink);background:var(--wash);padding:9px 14px;font-size:12px;min-height:44px}
.reset-search:hover,.filter-button:hover{background:#e3e9dc;color:var(--ink)}
main:focus{outline:none}
@media(max-width:760px){
 .menu-button{min-height:44px;padding:9px 13px}
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}

/* Small text, input zoom, and print legibility. */

.subtle-link{padding-block:4px;min-height:28px}
@media(max-width:760px){.search-field{font-size:16px;min-height:46px}.publication-row .publication-authors{font-size:12px}.publication-year{font-size:12px}.status{font-size:11px}}
@media print{.search-controls,.reset-search,.publication-arrow{display:none}.publication-row{grid-template-columns:55px 1fr;break-inside:avoid}.footer *{color:var(--ink)!important}.footer-bottom{border-color:var(--line)}.research-project,.tool-row,.contributor-list article{break-inside:avoid}}

.people-collaboration h2{font-size:25px;margin-bottom:13px}

/* One shared anchor offset avoids doubling the sticky-header allowance. */
.research-index{margin-top:18px;margin-bottom:20px;font-size:12px;gap:5px 18px}
.back-to-top{white-space:nowrap;text-underline-offset:4px}
.back-to-top:hover{text-decoration:underline}
@media print{.research-index,.back-to-top{display:none}}

/* Keep the footer return link usable by touch and visible as a link. */
.footer-bottom{align-items:center}
.back-to-top{display:inline-flex;align-items:center;min-height:28px;text-decoration:underline}
@media(max-width:760px){.footer-bottom{align-items:flex-start}.back-to-top{min-height:44px}}

.publication-fulltext{display:inline-block;min-height:24px;padding-block:2px;text-decoration:underline;text-underline-offset:3px;color:var(--ink);white-space:nowrap}

/* Keep every navigation link reachable in short landscape windows. */
@media(max-width:760px){.nav{max-height:calc(100vh - 79px);max-height:calc(100dvh - 79px);overflow-y:auto;overscroll-behavior-y:contain}}

/* Clear control boundaries and keyboard focus on both paper and dark surfaces. */
.feature-band .subtle-link{color:var(--accent)}
.search-field{border-color:var(--muted)}
.filter-button[aria-pressed=true]{border-color:var(--accent)}
.footer a:focus-visible{outline-color:#dce4da}

/* Let the intended image ratio size the mobile hero, and keep its action intact. */
.hero-figure img{height:auto}
@media(max-width:760px){.section-head .text-link{white-space:nowrap;flex-shrink:0}}

/* People uses the actual roster and biography without a placeholder illustration. */
.people-layout{display:block;max-width:800px}

.inline-link{text-decoration:underline;text-underline-offset:3px;color:var(--ink)}
.opportunities{padding-bottom:55px}.opportunity{display:grid;grid-template-columns:260px minmax(0,1fr);gap:40px;padding:30px 0;border-top:1px solid var(--line)}.opportunity h2{font-size:28px}.opportunity p{font-size:15px;line-height:1.85;max-width:760px}.opportunity .opportunity-deadline{font-size:13px;color:var(--ink)}
@media(max-width:760px){.opportunity{grid-template-columns:1fr;gap:16px;padding:26px 0}.opportunity p{font-size:14px}.opportunity h2{font-size:27px}.opportunities{padding-bottom:35px}}
@media print{.opportunity{break-inside:avoid}}

.citation{color:var(--accent);text-decoration:underline;text-underline-offset:3px;white-space:nowrap}

.student-projects-title{font-family:var(--serif);font-size:23px;margin-top:24px}.student-project-links{padding-left:19px;margin:12px 0;color:var(--ink);font-size:14px;line-height:1.9}.student-project-links a{text-decoration:underline;text-underline-offset:3px}.opportunity p.project-scope-note{font-size:13px;color:var(--muted)}
.student-project-page{padding-bottom:65px}.project-breadcrumb{display:inline-block;margin-top:30px;font-size:12px;text-decoration:underline;text-underline-offset:3px}.student-project-page .page-intro{padding-top:28px;padding-bottom:25px}.student-project-description{max-width:760px;font-size:15px;line-height:1.85}.student-project-work{margin-top:30px;max-width:760px}.student-project-work h2{font-size:27px}.student-project-work ul{padding-left:20px;margin-top:16px;font-size:15px;line-height:1.9}.student-project-page .project-links{margin-top:30px}
@media(max-width:760px){.student-project-description,.student-project-work ul{font-size:14px}.student-project-page{padding-bottom:35px}.student-project-page h1{font-size:34px}.student-projects-title{font-size:23px}.project-breadcrumb{margin-top:24px}}

.hero-figure{background:#203c34}.hero-figure img{height:420px;max-height:420px;aspect-ratio:auto;object-fit:contain;padding:16px}
.flower-viewer{margin-top:25px;background:var(--wash)}.model-frame{height:310px;position:relative;overflow:hidden}.model-frame canvas{display:block;width:100%;height:100%;outline-offset:-4px}.model-status{position:absolute;inset:0;display:flex;align-items:center;justify-content:center;font-size:12px;padding:20px}.flower-viewer figcaption{padding:12px;font-size:10px;line-height:1.8}.rotation-toggle{display:block;margin-top:8px;min-height:36px;padding:5px 10px;background:transparent;border:1px solid var(--muted);color:var(--ink);font-size:11px}
@media(max-width:760px){.hero-figure img{height:340px;max-height:340px}.model-frame{height:310px}.rotation-toggle{min-height:44px}}

/* Research reconstruction belongs with its project, with room for fine detail. */
.research-project .flower-viewer{margin:28px 0}.research-project .model-frame{height:440px}.research-project .flower-viewer figcaption{padding:14px 18px;display:flex;align-items:center;justify-content:space-between;gap:12px}.research-project .rotation-toggle{flex-shrink:0}
@media(max-width:760px){.research-project .model-frame{height:360px}.research-project .flower-viewer figcaption{padding:12px;flex-wrap:wrap}}

/* Focused readability pass: keep models full width and critical links legible. */
.research-project > p{max-width:72ch}
.project-links .text-link,.tool-row .text-link{font-size:12px;min-height:44px;align-items:center}
.hero-figure figcaption,.flower-viewer figcaption,.schematic figcaption,.research-photo figcaption{font-size:11px;line-height:1.7}
.opportunity-resources{display:flex;flex-wrap:wrap;align-items:center;gap:8px 24px;margin-top:12px;font-size:12px}
.opportunity-resources a{display:inline-flex;align-items:center;min-height:44px;color:var(--ink);text-decoration:underline;text-underline-offset:3px}
@media(max-width:760px){.project-links .text-link,.tool-row .text-link,.opportunity-resources{font-size:13px}}

/* Resolve inherited link sizing and the viewer's keyboard/flex states. */
.opportunity-resources a{font-size:inherit}
.research-project .rotation-toggle{margin-top:0}
.model-frame canvas:focus-visible{outline:2px solid var(--accent);outline-offset:-4px}
.schematic figcaption{color:var(--body)}
.flower-viewer .model-fallback{padding:24px 18px;font-size:13px;line-height:1.7}
.rotation-toggle{font-size:12px;min-height:44px}
@media(max-width:760px){.rotation-toggle{font-size:13px}.footer-links{font-size:13px}.footer-links a{display:inline-flex;align-items:center;min-height:44px}}

/* Give long paper titles and author lists the available width on phones. */
@media(max-width:480px){
 .publication-row{grid-template-columns:minmax(0,1fr) 16px;gap:8px 12px}
 .publication-year{grid-column:1/-1;padding-top:0}
 .publication-row>div{grid-column:1}
 .publication-arrow{grid-column:2;grid-row:2}
}

/* The lab's full name is identity, rather than fine-print metadata. */
.affiliation{font-size:12px}
@media(max-width:760px){.affiliation{font-size:11px}}

/* Let a loaded canvas shrink with its grid column during live resizing. */
.research-block > div,.model-frame{min-width:0}

/* A loaded reconstruction can be turned directly, without an extra label. */
.flower-viewer[data-loaded="true"] .model-frame canvas{cursor:grab}
.flower-viewer[data-loaded="true"] .model-frame canvas:active{cursor:grabbing}
.research-block .research-overview{min-width:0}

/* Keep the visible search cue above normal-text contrast across browsers. */
.search-field::placeholder{color:var(--muted);opacity:1}

/* Navigation remains usable until its mobile enhancement is ready. */
@media(max-width:760px){
 .masthead:not(.nav-enhanced){position:static}
 .masthead:not(.nav-enhanced) .masthead-inner{height:auto;min-height:79px;flex-wrap:wrap;padding-block:16px}
 .masthead:not(.nav-enhanced) .menu-button{display:none}
 .masthead:not(.nav-enhanced) .nav{display:flex;position:static;flex-direction:row;flex-wrap:wrap;padding:0;border:0;gap:4px 20px;width:100%;max-height:none;overflow:visible}
 .masthead:not(.nav-enhanced) .nav a{padding:8px 0}
}

/* A failed reconstruction keeps its message without an empty model viewport. */
.flower-viewer[data-loaded="error"] .model-frame{height:auto;min-height:112px}

.assembly-method{margin-top:27px}.assembly-method+.assembly-method{border-top:1px solid var(--line);padding-top:26px}.assembly-method h4{font-family:var(--serif);font-weight:400;color:var(--ink);font-size:24px;line-height:1.4;margin:10px 0 13px}.assembly-project>h3{margin-bottom:20px}

/* PI roles sit directly under the NYBG appointment; biography remains separate. */
.pi-appointments{list-style:none;padding:0;margin:8px 0 27px;font-size:13px;line-height:1.7;color:var(--muted)}.pi-appointments li+li{margin-top:5px}@media(max-width:760px){.pi-appointments{font-size:11px;margin-bottom:24px}}

/* A single current member has room for a concise introduction. */
.mentoring-section .current-member-list{display:block;max-width:740px}.current-member-list article{border-top:0;padding-top:0}.current-member-list h3{font-size:20px;line-height:1.5}.current-member-list p{font-size:14px;line-height:1.85}.current-member-list .member-role{font-size:12px;color:var(--muted);margin:5px 0 12px}@media(max-width:760px){.current-member-list h3{font-size:18px}.current-member-list p{font-size:13px}}

/* Modest fieldwork portrait beside the PI biography; no crop. */
.people-layout{display:grid;grid-template-columns:minmax(0,740px) 280px;gap:64px;max-width:none;align-items:start}.pi-fieldwork{margin:0;padding-top:6px}.pi-fieldwork img{width:100%;height:auto;display:block}@media(max-width:1050px){.people-layout{grid-template-columns:minmax(0,1fr) 240px;gap:36px}}@media(max-width:760px){.people-layout{grid-template-columns:1fr;gap:25px}.pi-fieldwork{order:-1;width:220px;max-width:100%;padding:0}.person-copy{padding-top:0}}

/* Keep appointments together, then align the fieldwork photograph with the biography. */
.people-layout{display:block;max-width:none}.pi-heading .pi-appointments{margin-bottom:0}.pi-bio-grid{display:grid;grid-template-columns:minmax(0,700px) 300px;gap:64px;align-items:start;margin-top:30px}.pi-bio{border:0;padding:0}.pi-fieldwork{width:300px;margin:0;padding:0}.pi-fieldwork figcaption{margin-top:10px;font:12px/1.6 var(--sans);color:var(--muted)}@media(max-width:1050px){.pi-bio-grid{grid-template-columns:minmax(0,1fr) 260px;gap:36px}.pi-fieldwork{width:260px}}@media(max-width:760px){.pi-bio-grid{display:flex;flex-direction:column;gap:25px;margin-top:24px}.pi-fieldwork{order:-1;width:260px;max-width:100%}.pi-heading{padding-top:24px}.pi-bio{padding:0}.pi-fieldwork figcaption{font-size:11px}}

/* Source-derived Pedicularis figure; intrinsic aspect ratio preserves the slide geometry. */
.pedicularis-phylogeny{margin:28px 0 30px}.pedicularis-phylogeny a{display:block;cursor:zoom-in}.pedicularis-phylogeny img{width:100%;height:auto}

/* Graduate directions foreground questions and independent research. */
.student-project-summary{display:block;font-size:13px;line-height:1.7;color:var(--muted);margin:4px 0 18px;max-width:66ch}
.student-project-questions{max-width:760px}.student-project-question{margin-top:32px}.student-project-question h2{font-family:var(--serif);font-size:27px;line-height:1.3;margin:0 0 16px;color:var(--ink)}.student-project-question p{font-size:15px;line-height:1.85;margin:0 0 16px}.student-project-description p{margin:0 0 16px}.student-project-page>.project-links{flex-wrap:wrap}
@media(max-width:760px){.student-project-question h2{font-size:25px}.student-project-question p{font-size:14px}.student-project-question{margin-top:28px}.student-project-summary{font-size:13px}}

/* Source-derived flower reconstruction loop. */
.atlas-media{margin:25px 0}
.atlas-video-frame{position:relative;background:#faf9f5;aspect-ratio:16/9}
.atlas-video-frame video,.atlas-video-frame picture,.atlas-video-frame img{position:absolute;inset:0;display:block;width:100%;height:100%;object-fit:contain}
.atlas-video-frame video{opacity:0}
.atlas-video-frame[data-ready="true"] video{opacity:1}
@media(max-width:600px){.atlas-video-frame{aspect-ratio:4/5}}


.student-project-bullets{margin:24px 0 0;padding-left:21px;font-size:15px;line-height:1.85}.student-project-bullets li{margin:0 0 12px;padding-left:4px}@media(max-width:760px){.student-project-bullets{font-size:14px}}

.student-project-support{max-width:760px;margin-top:28px;padding-top:18px;border-top:1px solid var(--line);font-size:13px;line-height:1.8;color:var(--muted)}

/* Keep the six-item primary navigation on one line at tablet widths. */
@media(min-width:761px) and (max-width:1050px){.nav{gap:15px}.nav a{white-space:nowrap}}

/* One consistent resource list on Join the Lab. */
.graduate-resources{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:0 24px;list-style:none;margin:24px 0 30px;padding:0}
.graduate-resources li{min-width:0}
.graduate-resources a{display:flex;align-items:center;justify-content:space-between;gap:16px;min-height:48px;padding:12px 0;border-bottom:1px solid var(--line);color:var(--ink);font-size:13px;font-weight:500;line-height:1.5;text-decoration:none}
.graduate-resources a span{flex-shrink:0;font-size:17px;font-weight:400}
.graduate-resources a:hover{border-color:var(--accent)}
@media(max-width:600px){.graduate-resources{grid-template-columns:1fr;gap:0}.graduate-resources a{min-height:48px}}

/* Full watercolor composition; title floats only where there is room. */
.hero--floral-atlas{grid-template-columns:1fr;gap:38px;padding-bottom:44px}
.hero--floral-atlas .hero-intro{display:grid;grid-template-columns:1.08fr 1fr;gap:70px;align-items:start}
.hero--floral-atlas .hero-deck{margin-top:0}
.hero--floral-atlas .hero-floral-atlas{background:var(--paper);overflow:visible}
.hero--floral-atlas .hero-floral-atlas img{width:100%;height:auto;max-height:none;aspect-ratio:auto;object-fit:contain;padding:0}
.hero--floral-atlas .hero-floral-atlas figcaption{position:static;display:block;max-width:none;margin-top:10px;padding:0;background:transparent;font-family:var(--sans);font-size:clamp(11px,.95vw,12px);line-height:1.5;letter-spacing:0;text-align:right;color:var(--muted)}
@media(max-width:1050px){.hero--floral-atlas .hero-intro{gap:38px}}
@media(max-width:760px){.hero--floral-atlas{gap:28px;padding-bottom:30px}.hero--floral-atlas .hero-intro{grid-template-columns:1fr;gap:22px}.hero--floral-atlas .hero-floral-atlas figcaption{position:static;max-width:none;margin-top:10px;font-size:12px;line-height:1.4}}

.publication-name-note{max-width:760px;margin-top:30px;font-size:12px;line-height:1.8;color:var(--muted)}